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Western Subterranean Termite
— Reticulitermes hesperus
Termites are small, white, tan, or black insects that can cause severe destruction to wooden structures. Termites belong to the insect order Isoptera, an ancient insect group that dates back more than 100 million years. The Latin name Isoptera means "equal wing"and refers to the fact that the front set of wings on a reproductive termite is similar in size and shape to the hind set.
Although many people think termites have only negative impacts, in nature they make many positive contributions to the world's ecosystems. Their greatest contribution is the role they play in recycling wood and plant material. Their tunneling efforts also help to ensure that soils are porous, contain nutrients, and are healthy enough to support plant growth. Termites are very important in the Sahara Desert where their activity helps to reclaim soils damaged by drying heat and wind and the overgrazing by livestock.
Termites become a problem when they consume structural lumber. Each year thousands of housing units in the United States require treatment for the control of termites. Termites may also damage utility poles and other wooden structures. Termite pests in California include drywood, dampwood, and subterranean species. These pests cause serious damage to wooden structures and posts and may also attack stored food, books, and household furniture.
IDENTIFICATION
Termites are social and can form large nests or colonies, consisting of very different looking individuals (castes). Physically the largest individual is the queen. Her function is to lay eggs, sometimes thousands in a single day. A king is always by her side. Other individuals have large heads with powerful jaws, or a bulblike head that squirts liquid. These individuals are called soldiers. But the largest group of termites in a colony is the workers. They toil long hours tending the queen, building the nest, or gathering food. While other species of social insects have workers, termites are unique among insects in that workers can be male or female. Surprisingly, termites can be long-lived: queens and kings can live for decades while individual workers can survive for several years.
Signs of termite infestation include swarming of winged forms in fall and spring and evidence of tunneling in wood. Darkening or blistering of wooden structural members is another indication of an infestation; wood in damaged areas is typically thin and easily punctured with a knife or screwdriver.
There are more than 2,500 different types of termites in the world and at least 17 different types of termites in California. However, most of this diversity can be lumped into four distinct groups: dampwood, drywood, subterranean, and mound builders. Mound builders do not occur in North America, but the other three species do. Dampwood termites are very limited in their distribution: most species are found only in California and the Pacific Northwest. Dampwood termites derive their name from the fact that they live and feed in very moist wood, especially in stumps and fallen trees on the forest floor. Drywood termites are common on most continents and can survive in very dry conditions, even in dead wood in deserts. They do not require contact with moisture or soil. Subterranean termites are very numerous in many parts of the world and live and breed in soil, sometimes many feet deep. Lastly, the mound builders are capable of building earthen towers 25 feet or more in height. Mounds may be located either in the soil or in trees, and where they occur in Africa, Australia, Southeast Asia, and parts of South America, they are very noticeable and remarkable.
Termites are sometimes confused with winged forms of ants, which also leave their underground nests in large numbers to establish new colonies and swarm in a manner similar to that of reproductive stages of termites. However, ants and termites can be distinguished by checking three features: antennae, wings, and waist.
Dampwood Termites
Dampwood termites are fairly common in central and northern coastal areas in California. They nest in wood buried in the ground, although contact with the ground is not necessary when infested wood is high in moisture. Because of their high moisture requirements, dampwood termites most often are found in cool, humid areas along the coast and are typical pests of beach houses. Winged reproductives typically swarm between July and October, but it is not unusual to see them at other times of the year. Dampwood termite winged reproductives (sometimes called swarmers) are attracted to lights.
Dampwood termites produce distinctive fecal pellets that are rounded at both ends, elongate, and lack the clear longitudinal ridges common to drywood termite pellets. Final confirmation of pellet identification may require help from an expert.
The Nevada dampwood termite, Zootermopsis nevadensis, occurs in the higher, drier mountainous areas of the Sierras where it is an occasional pest in mountain cabins and other forest structures; it also occurs along the northern California coast. The Pacific dampwood termite, Zootermopsis angusticollis, is almost one inch long, making it the largest of the termites occurring in California. Winged reproductives are dark brown with brown wings. Soldiers have a flattened brown or yellowish brown head with elongated black or dark brown mandibles. Nymphs are cream colored with a characteristic spotted abdominal pattern caused by food in their intestines. Nevada dampwood termites are slightly smaller and darker than the Pacific species; reproductives are about 3/4 inch long.
Drywood Termites
Drywood termites infest dry, undecayed wood, including structural lumber as well as dead limbs of native trees and shade and orchard trees, utility poles, posts, and lumber in storage. From these areas, winged reproductives seasonally migrate to nearby buildings and other structures usually on sunny days during fall months. Drywood termites are most prevalent in southern California (including the desert areas), but also occur along most coastal regions and in the Central Valley.
Drywood termites have a low moisture requirement and can tolerate dry conditions for prolonged periods. They remain entirely above ground and do not connect their nests to the soil. Piles of their fecal pellets, which are distinctive in appearance, may be a clue to their presence. The fecal pellets are elongate (about 3/100 inch long) with rounded ends and have six flattened or roundly depressed surfaces separated by six longitudinal ridges. They vary considerably in color, but appear granular and salt and pepperlike in color and appearance.
Winged adults of western drywood termites (Incisitermes minor) are dark brown with smoky black wings and have a reddish brown head and thorax; wing veins are black. These insects are noticeably larger than subterranean termites.
Subterranean Termites
Subterranean termites require moist environments. To satisfy this need, they usually nest in or near the soil and maintain some connection with the soil through tunnels in wood or through shelter tubes they construct. These shelter tubes are made of soil with bits of wood or even plasterboard (drywall). Much of the damage they cause occurs in foundation and structural support wood. Because of the moisture requirements of subterranean termites, they are often found in wood that has wood rot.
The western subterranean termite, Reticulitermes hesperus, is the most destructive termite found in California. Reproductive winged forms of subterranean termites are dark brown to brownish black, with brownish gray wings. On warm, sunny days following fall or sometimes spring rains, swarms of reproductives may be seen. Soldiers are wingless with white bodies and pale yellow heads. Their long, narrow heads have no eyes. Workers are slightly smaller than reproductives, wingless, and have a shorter head than soldiers; their color is similar to that of soldiers. In the desert areas of California, Heterotermes aureus, is the most destructive species of subterranean termites. Another destructive species in this group, the Formosan subterranean termite, Coptotermes formosanus, is now in California but restricted to a small area near San Diego. Unlike the western subterranean termite, Formosan subterranean termites swarm at dusk and are attracted to lights.
LIFE CYCLE
Most termite species swarm in late summer or fall, although spring swarms are not uncommon for subterranean and drywood termites. New kings and queens are winged during their early adult life but lose their wings after dispersing from their original colony. An infestation begins when a mated pair finds a suitable nesting site near or in wood and constructs a small chamber, which they enter and seal. Soon afterward, the female begins egg laying, and both the king and queen feed the young on predigested food until they are able to feed themselves. Most species of termites have microscopic, one-celled animals called protozoa within their intestines that help in converting wood (cellulose) into food for the colony.
Once workers and nymphs are produced, the king and queen are fed by the workers and cease feeding on wood. Termites go through incomplete metamorphosis with egg, nymph, and adult stages. Nymphs resemble adults but are smaller and are the most numerous stage in the colony. They also groom and feed one another and other colony members.
Information from UC Davis Agriculture and Natural Resources
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